Kungl. Vet.-Akad. Handlingar 7, 4: 263 (1929). ------- but:
K. svenska Vetensk-Akad. Handl., ser. 3 7(no. 4): 263 (1929) IF
Synonyms: Index Fungorum
Protolog: Data lacking.
Thallus: crustose, areolate to squamulose, dispersed or contiguous when dividing, up to 4 mm in diam. Areoles or squamules, irregular or round, up to 2 mm wide, up to 1-2 mm high, epruinose, becoming fissured and dividing, the rim down-turned or up-turned, quite polymorphous. The stipe can be quite thick and wider than half diameter of squamule, but at that width the squamule usually begins dividing. The upper surface, in a small population,can be very dark blackish-brown to a beautiful light brown. The lower surface is usually white and ecorticate. The cortex is usually + 70 µm thick, with thin syncortex which makes it usually look shiny. The algal layer is usually uneven, penetrated by hyphal bands. Apothecia one to several per areole or squamule, punctiform to 1 mm wide, the disk dark brown to reddish-brown, rough, sometimes pseudo-lecanorine and verruciform; often surrounded with thalline collar. Parathecium indistinct, to 10 µm wide. Hymenium usually 120-170 µm thick. Paraphyses 1-2 µm at base in diam., apices usually more closely septate in upper 30 µm, barely expanded to 2-2.5 µm, usually more closely septate in upper 30 µm. Epihymenium reddish-brown, 10 µm thick. Subhymenium 30-50 µm thick. Asci usually clavate, 80-125 x 25-35 µm. Ascospores 100 or more spores per ascus, 3-6 x 1-2 µm, usually narrowly ellipsoid, but quite various in size sometimes. Pycnidia are quite common, globose, 50 x 70 µm; conidia 3.0-4.5 x 1-2 µm (Knudsen 2008: 7).
Photobiont: Trebouxioid green alga.
Chemistry: No lichen substances. Spot tests negative. Secondary metabolites absent (Knudsen 2008: 8).
Ecology: On sandstone, granite, and volcanic rock from 500-3708 meters. A non-obligatory juvenile parasite on Aspicilia (Knudsen 2007).
Molecular data: Genbank
Distribution: Bolivia, California, Arizona. Database
Note:
Type locality: Data lacking.
Barcode: Data lacking.
Coordinates: Data lacking.