Folia Cryptog. Estonica 11: 1 (1979) ["1978"]. Type locality: Russia, Primorskiy Kray ["Far East, Primor'e Terr."], Sikhote-Alin Range, Mt. Chernyy, 1977, H. Trass Pri-563 (TU - holotype; CANL, H, NY, US - isotypes).
Type locality: Russia, Primorskiy Kray ["Far East, Primor'e Terr."], Sikhote-Alin Range, Mt. Chernyy, 1977, H. Trass Pri-563 (TU - holotype; CANL, H, NY, US - isotypes).