Bryologist 96: 80 (1993). Type locality: USA, California, Mendocino Co., Jackson State Forest, 1986, S. Hammer 1095 (FH - holotype; H, NY, SFSU, US, WIS - isotypes).
Type locality: USA, California, Mendocino Co., Jackson State Forest, 1986, S. Hammer 1095 (FH - holotype; H, NY, SFSU, US, WIS - isotypes).