Lecanora densa (Śliwa & Wetmore) Printzen

Bryologist 104, 3: 394 (2001)

Basionym: Lecanora varia subsp. densa Śliwa & Wetmore, Bryologist 103: 486 (2000).

 

Protologue: U.S.A. Colorado. Boulder Co., Green Mountain, 6.5 km (4 mi.) SW of Boulder, 2,200 m, abundant in self-pruned branches of crowded sapling of Pinus ponderosa, outer foothills of Front Range, 3.5.1977, Weber & Klockenbrink (Weber, Lich. exs. Colo 563)(MIN!, holotype).

Type locality: USA, Colorado, Boulder County, Green Mountain, 6.5 km (4 mi.) SW of Boulder, abundant in self-pruned branches of crowded sapling of Pinus ponderosa, outer foothills of Front Range, 2,200 m, 3.5.1977, Weber & Klockenbrink, Weber: Lich. Exs. Colo. 563 (MIN - holotype).

Description: Lichen. Thallus crustose, warted-areolate, sometimes endosubstratal and distinct only around apothecia, esorediate; areoles ± isodiametric in outline, sometimes incised or crenulate, av. diam. 0.15-0.25 mm, max. diam. 0.30-0.35 mm, weakly to moderately convex, surface yellowish beige to ochre, matt. Hypothallus ashgray to dark gray, between areoles. Apothecia rounded to deformed, rarely flexuose, single to densely crowded, sessile with constricted base, av. diam. 0.3-0.5 mm, max. diam. 0.6-0.8 mm; disc beige to orange-brown, often more or less olivaceous, flat to weakly, rarely moderately convex, whitish pruinose at least when wet, rarely epruinose, matt or slightly glossy; margin weakly to strongly prominent when young, persistent, more or less granular-knobby, matt, concolorous with thallus or slightly lighter. Amphithecium corticate, laterally 75-120 µm, basally 75-125 µm wide, nebula with irregularly entangled, short-celled hyphae with lumina ca 1.0-1.5(-2.0) µm wide, cortex 5-15 µm wide above, 25-65 µm near base, colorless within, outer part with granules like in epihymenium, of strongly gelatinized, anticlinally arranged hyphae with lumina of (1.0-)1.5-2.5 µm; parathecium 10-25 µm wide, or absent; hypothecium 40-60 µm high, colorless or pale yellowish; subhymenium 25-65 µm high; hymenium (35-)50-60 µm high colorless to pale yellowish; epihymenium 10-25 µm high, often streaking into hymenium, ochre to orange-brown, granular; paraphyses simple or weakly branched and anastomosing below, moderately branched above, with one µm, apically 1-2 µm wide lumina. Ascospores colorless, simple, (6.5-)8.0-9.5(-11.0) (5.0-)5.4-6.4(-7.5) µm. Pycnidia unknown (Printzen 2001: 394).

Photobiont: Data lacking.

Chemistry: Thallus C-, K-, P+; yellow; usnic acid, psoromic acid, ± 2´-O-demethylpsoromic acid.