Lichenologist 26, 1: 14 (1994). Type locality: Sweden, Västergötland, Otterstad, Kållandsö, Läckö peninsula opposite the castle, 1961, Degelius (UPS - holotype).
Type locality: Sweden, Västergötland, Otterstad, Kållandsö, Läckö peninsula opposite the castle, 1961, Degelius (UPS - holotype).